Has bestiality always been historically so taboo? Why? It doesn't result in pregnancy; did it result in diseases? Why did humans evolve to get so repulsed from touching genitals with other species?
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Possibly pedantic point that emphasises your hypothesis further -- in the version I read recently (New Jerusalem) Onan was engaging in 'coitus interruptus' / pulling out, rather than having a wank. May suggest more explicitly that the transgression was not fertilizing
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