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Any of these things can become a bottleneck. In fact, many unrelated species have developed forms of quasi-language, though nowhere near as advanced as that of humans, as they were bottlenecked by e.g. cognitive capabilities or environmental/social complexity.
This is also why the evolution of language is a very, very slow process. These factors are all interacting with each other, enabling or restricting the others -- a new factor becoming the new bottleneck whenever progress is made on one front. This is absolutely *co-evolution*.
A feedback loop that reinforces evolution. More complex language, changes environment and the agent too.
This is a great paper on the topic: https://cse.iitk.ac.in/users/hk/cs784/papers/comptalAndEvolAspectsOfLang.pdf …
see also, cell signalinghttps://en.wikipedia.org/wiki/Cell_signaling?wprov=sfti1 …
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