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The one consistent aspect of boi's use was that it had to do with atypical or countercultural masculinity. Black masculinity, skateboarders' masculinity, gay masculinity, or female masculinity. Well, it didn't just fall from there into the mainstream- it was pushed.
It was pushed by white racists using "digital blackface" to portray black people as negative stereotypes, with the hopes of turning white liberals against them.
And, it was pushed by white supremacists who fixated on a small controversy over whether the Dat Boi meme was culturally appropriative, seeing it as a challenge to take boi away from the communities where it was being used.
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