With EDNS0 enabled, the client can include a little fake sort of record that says "hey, I support EDNS0, and also I'm good if you send me large UDP responses ... up to 4,000 bytes, say".
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Because MTU discovery depends on state, and on ICMP messages being allowed, some folks do something like "MSS clamping" where for TCP they have the network actually meddle with the TCP connection (a little) to offer a different MSS to the other end.
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...and then HTTP/2 comes along and introduces its own flow control spec on top of TCP with zero guidance for implementation. Oh, and H/2 flow control is credit based whereas TCP's is not....
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