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Replying to @MorlockP
... says the man whose dog characters have a culture that leads them to auto-map where they pee. That sounds downstream of biology to me.
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Replying to @TheClarksTale
my point is that biology is downstream of culture too, and getting more so you think those dogs would be able to talk if humans hadn't had a culture that wanted talking dogs?
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Replying to @MorlockP
I get your point, but biology will always drive culture, though we're reaching the point where culture can also influence biology (arguably it has since we learned to make alcohol & began evolving to handle it).
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