Think of the peacock's tail, the most famous example, a hindrance for survival. It is said to have been selected because of its wastefulness, which would signal the vigor of its carrier.
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I tried to find where the actual criticism was in this very long paper, but lost patience.
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A signal need only be costly to be credible, not wasteful.
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The sentence towards the end starting with "Many studies refute the assumptions ..." is probably missing a bunch of words, or the word order is out of whack.
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